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Membracidae

Treehopper
Ceresa taurina.JPG
Ceresa taurina
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hemiptera
Suborder: Auchenorrhyncha
Infraorder: Cicadomorpha
Superfamily: Membracoidea
Family: Membracidae
Subfamilies

Centronodinae (disputed)
Centrotinae
Darninae
Endoiastinae
Heteronotinae
Membracinae
Nicomiinae (disputed)
Smiliinae
Stegaspidinae (disputed)
and see text

Synonyms

Nicomiidae


Centronodinae (disputed)
Centrotinae
Darninae
Endoiastinae
Heteronotinae
Membracinae
Nicomiinae (disputed)
Smiliinae
Stegaspidinae (disputed)
and see text

Nicomiidae

Treehoppers (more precisely typical treehoppers to distinguish them from the Aetalionidae) and thorn bugs are members of the family Membracidae, a group of insects related to the cicadas and the leafhoppers. About 3,200 species of treehoppers in over 400 genera are known. They are found on all continents except Antarctica; only three species are known from Europe. Individual treehoppers usually live for only a few months.

Thorn bugs, due to their unusual appearance, have long interested naturalists. They are best known for their enlarged and ornate pronotum, which most often resembles thorns, apparently to aid camouflage. In some species, the pronotum is a horn-like extension, but can form more bizarre shapes. The specialised pronotum (or helmet) may not be simply an expansion of the prothoracic sclerite, but a fused pair of dorsal appendages of the first thoracic segment.

These may be serial homologues of insect wings, which are dorsal appendages of the second and/or third thoracic segments. Evidence for this theory includes the development of the helmet, which arises as a pair of appendages attached to each side of the dorsal prothorax by an articulation with muscles and a flexible membrane that allow it to be mobile. Also, the same genes are involved in development of the helmet and the wings. Distinguishing males from females is accomplished only by looking at the genitalia.

Treehoppers pierce plant stems with their beaks, and feed upon sap. The immatures can frequently be found on herbaceous shrubs and grasses, whereas the adults more often frequent hardwood tree species. Excess sap becomes concentrated as honeydew, which often attracts ants. Some species have a well-developed ant mutualism, and these species are normally gregarious, as well, which attracts more ants. The ants provide protection from predators. Treehoppers mimic thorns to prevent predators from spotting them.


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Wikipedia

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